Note
Sperm nuclei are highly elongated and abruptly taper posteriorly. With the exception of one species, the nuclei lack a distinct centriolar fossa. Intracellular droplets resembling lipid extend from the extreme posterior region of the middlepiece to form a collar around the proximal portion of the axoneme. The presence of lipid like bodies in the middlepiece suggest that the sperm are long-lived and therefore require additional energy stores not found in most metazoan sperm.